List of butterflies of Sudan and South Sudan

This is a list of butterflies of Sudan and South Sudan. About 341 species are known from the Sudans,^[1] 6 of which are endemic.^[2]

Papilionidae

Papilioninae

Papilionini

• Papilio antimachus Drury, 1782
• Papilio zalmoxis Hewitson, 1864
• Papilio nireus lyaeus Doubleday, 1845
• Papilio nireus pseudonireus Felder & Felder, 1865
• Papilio plagiatus Aurivillius, 1898
• Papilio phorcas sudanicola Storace, 1965
• Papilio rex franciscae Carpenter, 1928
• Papilio mechowi Dewitz, 1881
• Papilio mechowianus Dewitz, 1885
• Papilio echerioides joiceyi Gabriel, 1945
• Papilio jacksoni imatonga Clifton & Collins, 1997
• Papilio nobilis didingensis Carpenter, 1928
• Papilio hesperus sudana Gabriel, 1945
• Papilio lormieri Distant, 1874
• Papilio mackinnoni Sharpe, 1891

Leptocercini

• Graphium antheus (Cramer, 1779)
• Graphium policenes policenes (Cramer, 1775)
• Graphium policenes telloi Hecq, 1999
• Graphium colonna (Ward, 1873)
• Graphium angolanus baronis (Ungemach, 1932)
• Graphium ridleyanus (White, 1843)
• Graphium leonidas (Fabricius, 1793)
• Graphium philonoe whalleyi (Talbot, 1929)
• Graphium almansor escherichi (Gaede, 1915)

Pieridae

Coliadinae

• Eurema brigitta (Stoll, [1780])
• Eurema hecabe solifera (Butler, 1875)
• Catopsilia florella (Fabricius, 1775)
• Colias electo pseudohecate Berger, 1940
• Colias marnoana Rogenhofer, 1884

Pierinae

• Colotis antevippe zera (Lucas, 1852)
• Colotis aurigineus (Butler, 1883)
• Colotis aurora evarne (Klug, 1829)
• Colotis celimene angusi Rothschild, 1921
• Colotis celimene sudanicus (Aurivillius, 1905)
• Colotis chrysonome (Klug, 1829)
• Colotis daira stygia (Felder & Felder, 1865)
• Colotis danae eupompe (Klug, 1829)
• Colotis elgonensis basilewskyi Berger, 1956
• Colotis ephyia (Klug, 1829)
• Colotis euippe exole (Reiche, 1850)
• Colotis halimede (Klug, 1829)
• Colotis hetaera aspasia (Ungemach, 1932)
• Colotis liagore (Klug, 1829)
• Colotis phisadia (Godart, 1819)
• Colotis pleione nilus Talbot, 1942
• Colotis protomedia (Klug, 1829)
• Colotis rogersi (Dixey, 1915)
• Colotis ungemachi (Le Cerf, 1922)
• Colotis vesta princeps Talbot, 1939
• Colotis vesta velleda (Lucas, 1852)
• Colotis vestalis castalis (Staudinger, 1884)
• Colotis eris (Klug, 1829)
• Calopieris eulimene (Klug, 1829)
• Eronia cleodora Hübner, 1823
• Eronia leda (Boisduval, 1847)
• Pinacopterix eriphia melanarge (Butler, 1886)
• Pinacopterix eriphia tritogenia (Klug, 1829)
• Nepheronia argia (Fabricius, 1775)
• Nepheronia buquetii (Boisduval, 1836)
• Nepheronia thalassina sinalata (Suffert, 1904)
• Euchloe falloui (Allard, 1867)
• Leptosia alcesta inalcesta Bernardi, 1959

Pierini

• Appias epaphia contracta (Butler, 1888)
• Appias phaola intermedia Dufrane, 1948
• Appias sabina (Felder & Felder, [1865])
• Appias sylvia sudanensis Talbot, 1932
• Pontia glauconome Klug, 1829
• Mylothris agathina (Cramer, 1779)
• Mylothris chloris (Fabricius, 1775)
• Mylothris jacksoni nagichota Talbot, 1944
• Mylothris rubricosta (Mabille, 1890)
• Mylothris rueppellii septentrionalis Carpenter, 1928
• Mylothris schumanni uniformis Talbot, 1944
• Dixeia dixeyi (Neave, 1904)
• Dixeia doxo doxo (Godart, 1819)
• Dixeia doxo venatus (Butler, 1871)
• Dixeia orbona vidua (Butler, 1900)
• Belenois aurota (Fabricius, 1793)
• Belenois creona (Cramer, [1776])
• Belenois raffrayi (Oberthür, 1878)
• Belenois solilucis loveni (Aurivillius, 1921)
• Belenois subeida (Felder & Felder, 1865)
• Belenois sudanensis (Talbot, 1929)
• Belenois theora laeta (Weymer, 1903)
• Belenois thysa meldolae Butler, 1872

Lycaenidae

Miletinae

Miletini

• Megalopalpus zymna (Westwood, 1851)
• Lachnocnema abyssinica Libert, 1996
• Lachnocnema emperamus (Snellen, 1872)
• Lachnocnema divergens Gaede, 1915
• Lachnocnema reutlingeri perspicua Libert, 1996

Poritiinae

Liptenini

• Alaena subrubra Bethune-Baker, 1915
• Pentila pauli multiplagata Bethune-Baker, 1908
• Pentila umangiana connectens Hulstaert, 1924
• Telipna sulpitia Hulstaert, 1924
• Telipna albofasciata laplumei Devos, 1917
• Ornipholidotos overlaeti intermedia Libert, 2005
• Ornipholidotos latimargo (Hawker-Smith, 1933)
• Mimacraea krausei karschioides Carpenter & Jackson, 1950
• Liptena ferrymani (Grose-Smith & Kirby, 1891)
• Liptena xanthostola xantha (Grose-Smith, 1901)
• Tetrarhanis ilala (Riley, 1929)
• Larinopoda lircaea (Hewitson, 1866)

Epitolini

• Stempfferia cercenoides (Holland, 1890)
• Cephetola dolorosa (Roche, 1954)
• Phytala elais ugandae Jackson, 1964

Aphnaeinae

• Pseudaletis agrippina Druce, 1888
• Chloroselas tamaniba (Walker, 1870)
• Chloroselas taposana Riley, 1932
• Cigaritis acamas bellatrix (Butler, 1886)
• Cigaritis nilus (Hewitson, 1865)
• Zeritis neriene Boisduval, 1836
• Zeritis pulcherrima Aurivillius, 1923
• Axiocerses harpax kadugli Talbot, 1935
• Axiocerses amanga (Westwood, 1881)
• Aphnaeus coronae Talbot, 1935

Theclinae

• Myrina silenus (Fabricius, 1775)
• Myrina subornata nuba Talbot, 1935
• Dapidodigma demeter sudsudana d'Abrera, 1980
• Hypolycaena liara Druce, 1890
• Hypolycaena obscura Stempffer, 1947
• Leptomyrina sudanica Stempffer, 1964
• Iolaus alienus ugandae Stempffer, 1953
• Iolaus scintillans Aurivillius, 1905
• Iolaus sudanicus Aurivillius, 1905
• Iolaus pallene (Wallengren, 1857)
• Iolaus menas Druce, 1890
• Iolaus ismenias (Klug, 1834)
• Stugeta marmoreus (Butler, 1866)
• Paradeudorix ituri ugandae (Talbot, 1935)
• Hypomyrina mimetica Libert, 2004
• Deudorix dinochares Grose-Smith, 1887
• Deudorix dinomenes diomedes Jackson, 1966
• Deudorix livia (Klug, 1834)
• Deudorix lorisona baronica Ungemach, 1932
• Capys bamptoni Henning & Henning, 1988

Polyommatinae

Lycaenesthini

• Anthene butleri (Oberthür, 1880)
• Anthene crawshayi minuta (Bethune-Baker, 1916)
• Anthene hodsoni (Talbot, 1935)
• Anthene lunulata (Trimen, 1894)
• Anthene wilsoni (Talbot, 1935)
• Anthene lusones (Hewitson, 1874)
• Anthene nigeriae (Aurivillius, 1905)

Polyommatini

• Cupidopsis jobates mauritanica Riley, 1932
• Uranothauma delatorum Heron, 1909
• Uranothauma heritsia intermedia (Tite, 1958)
• Leptotes marginalis (Stempffer, 1944)
• Tuxentius margaritaceus (Sharpe, 1892)
• Tarucus balkanicus (Freyer, 1843)
• Tarucus legrasi Stempffer, 1948
• Tarucus rosacea (Austaut, 1885)
• Tarucus theophrastus (Fabricius, 1793)
• Tarucus ungemachi Stempffer, 1942
• Zizeeria karsandra (Moore, 1865)
• Actizera stellata (Trimen, 1883)
• Euchrysops alberta (Butler, 1901)
• Euchrysops albistriata (Capronnier, 1889)
• Euchrysops crawshayi (Butler, 1899)
• Euchrysops cyclopteris (Butler, 1876)
• Euchrysops nilotica (Aurivillius, 1904)
• Chilades eleusis (Demaison, 1888)
• Lepidochrysops albilinea Tite, 1959
• Lepidochrysops negus wau (Wichgraf, 1921)
• Lepidochrysops nigrita Tite, 1959
• Lepidochrysops polydialecta (Bethune-Baker, [1923])
• Lepidochrysops victoriae occidentalis Libert & Collins, 2001

Nymphalidae

Danainae

Danaini

• Danaus chrysippus chrysippus (Linnaeus, 1758)
• Danaus chrysippus alcippus (Cramer, 1777)
• Danaus dorippus (Klug, 1845)
• Tirumala formosa neumanni (Rothschild, 1902)
• Tirumala petiverana (Doubleday, 1847)
• Amauris niavius aethiops Rothschild & Jordan, 1903
• Amauris tartarea Mabille, 1876
• Amauris albimaculata sudanica Talbot, 1940
• Amauris echeria mongallensis Carpenter, 1928
• Amauris echeria steckeri Kheil, 1890
• Amauris hecate (Butler, 1866)

Satyrinae

Melanitini

• Gnophodes betsimena parmeno Doubleday, 1849
• Melanitis libya Distant, 1882

Satyrini

• Bicyclus angulosa (Butler, 1868)
• Bicyclus campus (Karsch, 1893)
• Bicyclus istaris (Plötz, 1880)
• Bicyclus kenia (Rogenhofer, 1891)
• Bicyclus milyas (Hewitson, 1864)
• Bicyclus pavonis (Butler, 1876)
• Bicyclus martius sanaos (Hewitson, 1866)
• Bicyclus saussurei angustus Condamin, 1970
• Heteropsis perspicua (Trimen, 1873)
• Ypthima albida Butler, 1888
• Ypthima antennata van Son, 1955
• Ypthima condamini Kielland, 1982
• Ypthima doleta Kirby, 1880
• Ypthima pupillaris Butler, 1888
• Ypthima simplicia Butler, 1876
• Ypthimomorpha itonia (Hewitson, 1865)

Charaxinae

Charaxini

• Charaxes varanes vologeses (Mabille, 1876)
• Charaxes fulvescens monitor Rothschild, 1900
• Charaxes boueti rectans Rothschild & Jordan, 1903
• Charaxes cynthia kinduana Le Cerf, 1923
• Charaxes lactetinctus ungemachi Le Cerf, 1927
• Charaxes jasius Poulton, 1926
• Charaxes epijasius Reiche, 1850
• Charaxes hansali baringana Rothschild, 1905
• Charaxes brutus angustus Rothschild, 1900
• Charaxes julius somalicus Rothschild, 1900
• Charaxes ansorgei kinyeti Plantrou, 1989
• Charaxes pollux (Cramer, 1775)
• Charaxes druceanus vivianae Plantrou, 1982
• Charaxes eudoxus imatongensis Plantrou, 1982
• Charaxes eudoxus mechowi Rothschild, 1900
• Charaxes numenes aequatorialis van Someren, 1972
• Charaxes tiridates tiridatinus Röber, 1936
• Charaxes bipunctatus ugandensis van Someren, 1972
• Charaxes smaragdalis gobyae Plantrou, 1989
• Charaxes etesipe (Godart, 1824)
• Charaxes achaemenes monticola van Someren, 1970
• Charaxes jahlusa ganalensis Carpenter, 1937
• Charaxes eupale latimargo Joicey & Talbot, 1921
• Charaxes dilutus ngonga van Someren, 1974
• Charaxes baumanni didingensis van Someren, 1971
• Charaxes etheocles carpenteri van Someren & Jackson, 1957
• Charaxes cedreatis Hewitson, 1874
• Charaxes viola picta van Someren & Jackson, 1952
• Charaxes paphianus subpallida Joicey & Talbot, 1925
• Charaxes kahldeni Homeyer & Dewitz, 1882
• Charaxes zoolina (Westwood, [1850])
• Charaxes lycurgus bernardiana Plantrou, 1978
• Charaxes zelica depuncta Joicey & Talbot, 1921
• Charaxes amandae Rydon, 1989

Euxanthini

• Charaxes crossleyi ansorgei (Rothschild, 1903)

Apaturinae

• Apaturopsis cleochares (Hewitson, 1873)

Nymphalinae

Nymphalini

• Vanessa dimorphica (Howarth, 1966)
• Junonia chorimene (Guérin-Méneville, 1844)
• Junonia terea tereoides (Butler, 1901)
• Junonia westermanni Westwood, 1870
• Protogoniomorpha parhassus (Drury, 1782)
• Protogoniomorpha temora (Felder & Felder, 1867)
• Precis archesia ugandensis (McLeod, 1980)
• Precis coelestina Dewitz, 1879
• Precis octavia (Cramer, 1777)
• Precis tugela aurorina (Butler, 1894)
• Hypolimnas anthedon (Doubleday, 1845)
• Hypolimnas misippus (Linnaeus, 1764)
• Hypolimnas salmacis platydema Rothschild & Jordan, 1903
• Mallika jacksoni (Sharpe, 1896)

Biblidinae

Biblidini

• Byblia anvatara acheloia (Wallengren, 1857)
• Mesoxantha ethosea reducta Rothschild, 1918
• Neptidopsis ophione nucleata Grünberg, 1911

Limenitinae

Limenitidini

• Pseudoneptis bugandensis Stoneham, 1935
• Pseudacraea acholica Riley, 1932
• Pseudacraea boisduvalii (Doubleday, 1845)
• Pseudacraea eurytus (Linnaeus, 1758)
• Pseudacraea lucretia protracta (Butler, 1874)

Neptidini

• Neptis metella (Doubleday, 1848)
• Neptis nemetes nemetes Hewitson, 1868
• Neptis nemetes margueriteae Fox, 1968
• Neptis occidentalis Rothschild, 1918

Adoliadini

• Catuna crithea (Drury, 1773)
• Pseudargynnis hegemone (Godart, 1819)
• Euphaedra medon fraudata van Someren, 1935
• Euphaedra neumanni Rothschild, 1902
• Euphaedra harpalyce sudanensis Talbot, 1929

Heliconiinae

Acraeini

• Acraea chilo Godman, 1880
• Acraea insignis Distant, 1880
• Acraea neobule Doubleday, 1847
• Acraea pseudolycia astrigera Butler, 1899
• Acraea quirina (Fabricius, 1781)
• Acraea zetes (Linnaeus, 1758)
• Acraea abdera Hewitson, 1852
• Acraea cepheus (Linnaeus, 1758)
• Acraea egina (Cramer, 1775)
• Acraea guluensis Le Doux, 1932
• Acraea caecilia (Fabricius, 1781)
• Acraea caldarena Hewitson, 1877
• Acraea doubledayi Guérin-Méneville, 1849
• Acraea marnois Rogenhofer, 1890
• Acraea pseudegina Westwood, 1852
• Acraea sykesi Sharpe, 1902
• Acraea aganice orientalis (Ungemach, 1932)
• Acraea epaea paragea (Grose-Smith, 1900)
• Acraea epiprotea (Butler, 1874)
• Acraea macarista (Sharpe, 1906)
• Acraea poggei nelsoni Grose-Smith & Kirby, 1892
• Acraea pseuderyta Godman & Salvin, 1890
• Acraea acerata Hewitson, 1874
• Acraea bonasia (Fabricius, 1775)
• Acraea encedana Pierre, 1976
• Acraea serena (Fabricius, 1775)
• Acraea esebria Hewitson, 1861
• Acraea jodutta (Fabricius, 1793)
• Acraea johnstoni Godman, 1885
• Acraea lycoa Godart, 1819
• Acraea pharsalus pharsalus Ward, 1871
• Acraea pharsalus rhodina Rothschild, 1902
• Acraea oreas Sharpe, 1891

Argynnini

• Issoria hanningtoni imatonga (Riley, 1932)

Vagrantini

• Lachnoptera anticlia (Hübner, 1819)
• Phalanta eurytis eurytis (Doubleday, 1847)
• Phalanta eurytis microps (Rothschild & Jordan, 1903)
• Phalanta phalantha aethiopica (Rothschild & Jordan, 1903)

Hesperiidae

Coeliadinae

• Coeliades forestan (Stoll, [1782])
• Coeliades keithloa menelik (Ungemach, 1932)
• Coeliades pisistratus (Fabricius, 1793)

Pyrginae

Celaenorrhinini

• Celaenorrhinus proxima (Mabille, 1877)
• Eretis lugens (Rogenhofer, 1891)
• Eretis melania Mabille, 1891
• Eretis mixta Evans, 1937
• Sarangesa aza Evans, 1951
• Sarangesa brigida sanaga Miller, 1964
• Sarangesa laelius (Mabille, 1877)
• Sarangesa lucidella (Mabille, 1891)
• Sarangesa phidyle (Walker, 1870)

Tagiadini

• Tagiades flesus (Fabricius, 1781)
• Eagris decastigma purpura Evans, 1937
• Eagris denuba obliterata Carpenter, 1928
• Eagris lucetia (Hewitson, 1875)
• Eagris tigris liberti Collins & Larsen, 2005
• Caprona adelica Karsch, 1892
• Netrobalane canopus (Trimen, 1864)
• Abantis bismarcki Karsch, 1892

Carcharodini

• Spialia doris (Walker, 1870)
• Spialia zebra bifida (Higgins, 1924)

Hesperiinae

Aeromachini

• Prosopalpus styla Evans, 1937
• Kedestes protensa Butler, 1901
• Kedestes rogersi Druce, 1907
• Gorgyra aretina (Hewitson, 1878)
• Pardaleodes incerta (Snellen, 1872)
• Pardaleodes sator pusiella Mabille, 1877
• Pardaleodes tibullus (Fabricius, 1793)
• Osmodes thora (Plötz, 1884)
• Acleros mackenii olaus (Plötz, 1884)
• Andronymus caesar philander (Hopffer, 1855)
• Chondrolepis niveicornis (Plötz, 1883)
• Zophopetes dysmephila (Trimen, 1868)
• Gretna cylinda (Hewitson, 1876)
• Caenides dacena (Hewitson, 1876)
• Fresna nyassae (Hewitson, 1878)

Baorini

• Borbo fallax (Gaede, 1916)
• Borbo perobscura (Druce, 1912)
• Gegenes nostrodamus (Fabricius, 1793)

Heteropterinae

• Metisella quadrisignatus (Butler, 1894)
• Metisella tsadicus (Aurivillius, 1905)
• Lepella lepeletier (Latreille, 1824)

References

• Seitz, A. Die Gross-Schmetterlinge der Erde 13: Die Afrikanischen Tagfalter. Plates
• Seitz, A. Die Gross-Schmetterlinge der Erde 13: Die Afrikanischen Tagfalter. Text (in German)